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@media(max-width:1350px){.brand small{display:none}}
@media(max-width:1180px){.header{height:80px}.menu-toggle{display:block}.header nav{display:none;position:absolute;left:0;right:0;top:80px;background:white;padding:22px 25px 27px;flex-direction:column;align-items:stretch;gap:0;box-shadow:0 12px 25px #102d201c}.header nav.open{display:flex}.header nav a{padding:12px 0}.header nav .nav-contact{text-align:center;margin-top:10px}.bio-layout{grid-template-columns:210px minmax(0,1fr);gap:45px}.media-grid{gap:24px}}
@media(max-width:900px){.media-grid{grid-template-columns:1fr}.media-card{padding:25px 0 10px}.media-card h3{font-size:43px;margin:20px 0}.media-card p{max-width:650px;margin-bottom:22px}.bio-layout{display:block}.bio-index{position:static;display:flex;flex-flow:row wrap;gap:6px 22px;margin-bottom:65px}.bio-index .kicker{width:100%;margin-bottom:10px}.bio-index a{padding:10px 0}.bio-hero{grid-template-columns:55% 45%}.bio-hero-copy{padding-right:30px}}
@media(max-width:720px){.authority-strip{padding:23px 24px;gap:18px;grid-template-columns:1fr}.authority-strip a,.authority-strip a:first-child{display:flex;flex-direction:row;align-items:center;justify-content:space-between;padding:12px 0;border-left:0;border-bottom:1px solid #a8bd95}.authority-strip a:last-child{border-bottom:0}.authority-strip span{font-size:11px;letter-spacing:.07em}.authority-strip strong{font-size:24px}.hero-copy{padding-top:48px}.hero .eyebrow{font-size:12px;letter-spacing:.07em;gap:10px}.hero h1{font-size:clamp(43px,9.7vw,63px)}.hero-actions{gap:25px}.about .button{font-size:12px;gap:14px;padding:20px 18px}.book-feature{grid-template-columns:1fr}.book-feature .cover{height:410px}.book-feature .book-info{padding:28px 25px}.book-feature .book-info h3{font-size:36px}.book .cover{height:300px;padding:23px 14px}.book-feature .cover{height:410px;padding:30px}.book-info{padding:21px 18px;min-height:300px}.book-info p{font-size:16px}.book-info h3{font-size:23px}.book-info a{font-size:13px}.bio-hero{display:flex;flex-direction:column}.bio-hero-copy{padding:28px 24px 50px}.back-link{margin-bottom:40px}.bio-hero .kicker{font-size:12px}.bio-hero h1{font-size:clamp(60px,15vw,94px)}.bio-portrait{min-height:430px;max-height:540px}.bio-portrait img{object-position:center 35%}.bio-layout{padding:60px 24px}.bio-index{margin-bottom:55px;gap:4px 20px}.bio-story p{font-size:18px;line-height:1.75}.bio-story .standfirst{font-size:22px}.bio-story section{margin-bottom:46px;padding-bottom:42px}.bio-story h2{font-size:35px}.research-reference{padding:24px 21px}.research-reference h3{font-size:20px;line-height:1.45}.bio-contact{padding:65px 24px}.book-source-note{line-height:1.7}}
@media(max-width:520px){.books-grid{grid-template-columns:1fr}.book .cover{height:370px;padding:26px}.book-info{min-height:240px;padding:25px}.book-feature .cover{height:425px;padding:30px}.authority-strip strong{font-size:22px}.authority-strip small{margin-left:4px}}

/* Thematic pages and mobile navigation. */
html{scroll-padding-top:110px}body{overflow-wrap:break-word}.header{position:sticky;top:0;border-bottom:1px solid #e7ebe5}.header nav a{min-height:44px;display:flex;align-items:center;justify-content:center}.header nav .nav-contact{padding:13px 18px}.menu-toggle{min-height:44px;min-width:80px}.button,.text-link{min-height:44px;line-height:1.5}.hero-image img{object-position:50% 42%}.hero{grid-template-columns:55% 45%}.hero-copy{padding-right:5vw}.hero-index{font-size:12px}.page-intro{padding:45px max(4.2vw,24px) 75px;background:var(--pine);color:white;display:flex;flex-direction:column;align-items:flex-start}.page-intro .back-link{margin-bottom:48px}.page-intro .kicker{color:var(--lime)}.page-intro h1{font-size:clamp(47px,6vw,86px);line-height:1.06;margin:26px 0;max-width:1000px}.page-intro p{font-size:21px;line-height:1.6;color:#d5e2d1;max-width:740px;margin:0}.page-intro em{font-style:normal;color:#adc497}.catalog-page .books>.section-heading,.photos-page .fungi>.section-heading,.events-page .events .section-intro>h2{display:none}.catalog-page .books{padding-top:70px}.events-page .events{padding-top:75px;padding-bottom:75px}.events-page .event-photo{margin-top:20px;max-width:none}.events-page .events-content{padding-top:20px}.events-page .event-photo{aspect-ratio:auto}.events-page .event-photo img{height:auto;object-fit:contain}.event-album{padding-top:80px}.event-gallery{display:grid;grid-template-columns:1fr 1fr;gap:48px 30px;align-items:start}.photo-story{margin:0;min-width:0}.photo-story:first-child{grid-column:1/-1}.photo-story a{display:block;background:#edf0e9}.photo-story img{width:100%;height:auto;object-fit:contain}.photo-story:first-child img{max-height:760px}.photo-story figcaption,.field-photo figcaption{font-size:15px;line-height:1.65;color:#5b6c60;padding-top:13px}.photo-story a:hover img{opacity:.95}.photos-page .gallery{grid-template-columns:repeat(3,minmax(0,1fr))}.photos-page .gallery 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.standfirst{font-size:25px;color:var(--ink)}.home-book .button{margin-top:20px}.explore{display:grid;grid-template-columns:1fr 1.15fr;gap:9vw}.explore h2{font-size:clamp(38px,3.6vw,57px);margin:27px 0 0}.explore-links{border-top:2px solid var(--ink)}.explore-links a{display:grid;grid-template-columns:1fr auto;gap:12px;text-decoration:none;padding:25px 0;border-bottom:1px solid var(--line)}.explore-links span{font:800 13px Manrope,sans-serif;letter-spacing:.09em;color:#567148;grid-column:1}.explore-links strong{font:600 23px/1.4 Manrope,sans-serif;letter-spacing:-.03em;grid-column:1}.explore-links b{grid-column:2;grid-row:1/3;align-self:center;font-size:27px}.explore-links a:hover strong{color:#567148}.contact-layout{display:grid;grid-template-columns:1.2fr 1fr;gap:7vw}.contact-layout .contact-mail{font-size:clamp(23px,3.4vw,48px);display:flex;margin:26px 0 36px;letter-spacing:-.045em;width:fit-content;max-width:100%}.contact-layout p{font-size:19px;line-height:1.75;color:#4e6253;max-width:590px}.contact-layout .contact-privacy{font-size:14px}.contact-areas{border-top:2px solid var(--ink);padding-top:28px}.contact-areas h2{font-size:clamp(28px,2.5vw,40px);line-height:1.3;letter-spacing:-.04em;margin-bottom:30px}.contact-areas p{font-size:17px;border-bottom:1px solid var(--line);padding-bottom:15px}.legal-layout{padding-top:65px}.legal-copy{max-width:880px;margin:0 auto}.legal-copy h2{font-size:29px;line-height:1.3;letter-spacing:-.035em;margin:42px 0 20px}.legal-copy p{font-size:18px;line-height:1.8;color:#43574a}.legal-copy a{text-underline-offset:4px;overflow-wrap:anywhere}.privacy-scope{padding:26px 30px;background:#edf2e8;border-left:3px solid #72945d}.privacy-scope h2{margin-top:0}.privacy-scope p{margin-bottom:0}.legal-references{border-top:1px solid var(--line);margin-top:40px;padding-top:25px;display:flex;flex-direction:column;gap:20px}.legal-references a{font-size:16px}.footer-email{font-size:15px;text-underline-offset:4px;color:#d2decb;margin-top:9px}footer>div:nth-child(2){align-content:flex-start;gap:10px 25px}footer>div:nth-child(2) a{font-size:13px;min-height:44px;display:flex;align-items:center}.contact .kicker{font-size:13px}.contact p{line-height:1.7}
@media(max-width:1180px){.header nav{max-height:calc(100dvh - 80px);overflow-y:auto;overscroll-behavior:contain}.header nav a{font-size:16px;justify-content:flex-start;padding:13px 0;min-height:49px}.header nav .nav-contact{justify-content:center}.bio-index{top:108px}}
@media(max-width:900px){.home-event{grid-template-columns:1fr}.home-event-copy{padding:60px max(4.2vw,24px)}.home-event-image img{max-height:650px}.contact-layout{grid-template-columns:1fr;gap:55px}.field-photo{grid-template-columns:1fr;gap:40px}.bio-index{position:static}.home-book{gap:45px}.explore{grid-template-columns:1fr;gap:40px}}
@media(max-width:720px){.hero{display:flex;flex-direction:column}.hero-copy{padding:42px 24px 40px}.hero-image{min-height:0;aspect-ratio:4/5}.hero-image img{object-position:50% 42%}.hero h1{font-size:clamp(37px,9.4vw,61px)}.hero .lead{font-size:18px;line-height:1.7}.hero .eyebrow{font-size:11px;letter-spacing:.07em}.hero-actions{gap:18px;margin:12px 0 32px}.hero-actions .button{font-size:13px;padding:18px 19px;gap:15px}.hero-actions .text-link{font-size:13px}.hero-index{font-size:11px}.page-intro{padding:30px 24px 53px}.page-intro .back-link{margin-bottom:32px}.page-intro h1{font-size:clamp(41px,10vw,65px);margin:23px 0}.page-intro p{font-size:19px}.catalog-page .books{padding-top:45px}.event-gallery{grid-template-columns:1fr;gap:32px}.photo-story:first-child{grid-column:auto}.events-page .events{padding:50px 24px}.event-album{padding-top:55px}.photos-page .gallery{grid-template-columns:1fr;gap:22px}.photos-page .gallery figure{aspect-ratio:auto}.photos-page .gallery img{height:auto}.home-book{grid-template-columns:1fr;gap:40px}.home-book-cover{height:360px}.home-book h2{font-size:48px}.home-book p{font-size:18px}.home-book .button{font-size:12px;letter-spacing:.02em;padding:18px 17px}.home-event h2{font-size:47px}.home-event p{font-size:18px}.home-event .text-link{font-size:13px;line-height:1.7}.explore h2{font-size:39px}.explore-links strong{font-size:22px}.contact-layout .contact-mail{font-size:clamp(22px,6.6vw,36px);line-height:1.5;letter-spacing:-.04em}.contact-areas h2{font-size:31px}.legal-layout{padding-top:40px}.legal-copy h2{font-size:26px}.legal-copy p{font-size:17px}.privacy-scope{padding:22px}.video-label h3{font-size:24px}.bio-portrait{min-height:0;aspect-ratio:1/1;max-height:none}.bio-portrait img{object-position:center}.bio-contact>a{font-size:clamp(21px,5.9vw,37px)}}
@media(max-width:380px){.hero .eyebrow{font-size:10px;letter-spacing:.045em}.hero .rule{width:22px}.authority-strip span{font-size:10px}.authority-strip strong{font-size:20px}.contact-mail{font-size:23px}.about .button{font-size:11px}.page-intro h1{font-size:40px}}
.events-page .events .section-intro>h2{display:block;font-size:clamp(38px,3.6vw,58px);margin:28px 0}.events-intro-text{font-size:19px;line-height:1.7;color:#526158;max-width:570px}.button{max-width:100%;white-space:normal}.header nav a{overflow-wrap:anywhere}.photos-page .gallery a{display:block;height:100%}.footer-email{max-width:100%;overflow-wrap:anywhere}@media(max-width:720px){.photos-page .gallery a{height:auto}.events-intro-text{font-size:18px}}
footer{grid-template-columns:minmax(0,1fr) minmax(0,1.7fr)}footer>div:nth-child(2){flex-wrap:wrap;justify-content:flex-end}.home-page .hero-copy,.home-page .hero-image,.bio-hero-copy{min-width:0}@media(max-width:720px){footer>div:nth-child(2){justify-content:flex-start}}

/* Photograph albums: preserve original framing at every screen size. */
.photos-page .fungi>.section-heading{display:flex;margin-bottom:36px}
.photos-page .fungi>.section-heading h2,.field-album .section-heading h2{font-size:clamp(36px,4vw,58px);margin-top:22px}
.field-album{padding-top:0}
.field-gallery{columns:2;column-gap:24px}
.field-gallery figure{margin:0 0 24px;break-inside:avoid}
.field-gallery a{display:block;background:var(--cream)}
.field-gallery img{display:block;width:100%;height:auto}
.field-gallery a:focus-visible,.photos-page .gallery a:focus-visible{outline:3px solid #557565;outline-offset:5px}
.gallery-more{margin-top:28px}
@media(max-width:720px){.photos-page .fungi>.section-heading{display:block}.field-gallery{columns:1}.field-gallery figure{margin-bottom:22px}}

.home-page .hero-image img{object-position:60% 34%}@media(max-width:720px){.home-page .hero-image{aspect-ratio:1}.home-page .hero-image img{object-position:center}}

/* Natural history photographs: compact thumbnails, intact originals. */
.photos-page .page-intro{padding-bottom:42px}
.photos-page .page-intro .back-link{margin-bottom:24px}
.photo-collection{padding-top:38px}
.nature-gallery{display:grid;grid-template-columns:repeat(5,minmax(0,1fr));gap:14px}
.nature-gallery figure{margin:0;min-width:0;scroll-margin-top:12px}
.nature-gallery a,.home-photo-grid a{display:block;aspect-ratio:1;background:#ecf0e9;cursor:zoom-in}
.nature-gallery img,.home-photo-grid img{width:100%;height:100%;object-fit:contain}
.nature-gallery a:focus-visible,.home-photo-grid a:focus-visible{outline:3px solid #557565;outline-offset:4px}
.nature-gallery figure:target{outline:3px solid #557565;outline-offset:4px}
.nature-gallery a:hover img,.home-photo-grid a:hover img{opacity:.9}
.home-photographs{padding-top:70px;padding-bottom:75px;background:var(--cream)}
.home-photographs .section-heading{margin-bottom:32px;align-items:center}
.home-photographs h2{font-size:clamp(34px,3.5vw,54px);margin-top:22px}
.home-photo-grid{display:grid;grid-template-columns:repeat(6,minmax(0,1fr));gap:12px}
.home-photo-grid a{cursor:pointer}
.photo-open{overflow:hidden}
.photo-viewer{width:min(1120px,96vw);max-width:96vw;max-height:94dvh;padding:0;border:0;background:#10241c;color:#fff;overflow:auto}
.photo-viewer::backdrop{background:#06110be8}
.viewer-panel{padding:18px 22px 20px}
.viewer-header{display:flex;align-items:center;gap:20px;margin-bottom:12px;font-size:14px}
#photo-viewer-title{font-weight:700;margin-right:auto}
#photo-counter{font-variant-numeric:tabular-nums;white-space:nowrap;color:#cdddc7}
.photo-viewer button{border:1px solid #788f7d;color:#fff;background:#243e30;min-width:48px;min-height:48px;cursor:pointer;font-size:25px}
.photo-viewer button:hover{background:#3a563e}
.photo-viewer button:focus-visible{outline:3px solid #d9e7b9;outline-offset:3px}
.photo-viewer .viewer-close{font-size:13px;font-weight:700;padding:0 15px}
.viewer-stage{display:grid;grid-template-columns:48px minmax(0,1fr) 48px;align-items:center;gap:16px}
#photo-full{width:100%;height:70dvh;max-height:760px;object-fit:contain;background:#0c1a14}
#photo-caption{font-size:15px;line-height:1.6;margin:15px auto 0;text-align:center;color:#d7e3d2;max-width:850px}
@media(min-width:1350px){.nature-gallery{grid-template-columns:repeat(6,minmax(0,1fr))}}
@media(max-width:1000px){.nature-gallery{grid-template-columns:repeat(4,minmax(0,1fr))}}
@media(max-width:720px){.nature-gallery{grid-template-columns:repeat(3,minmax(0,1fr));gap:10px}.home-photo-grid{grid-template-columns:repeat(3,minmax(0,1fr));gap:10px}.home-photographs{padding-top:50px;padding-bottom:55px}.home-photographs .section-heading{display:block}.home-photographs .text-link{margin-top:20px}.viewer-panel{padding:12px}.viewer-header{gap:12px;flex-wrap:wrap}.viewer-stage{grid-template-columns:48px minmax(0,1fr) 48px;gap:6px}.viewer-stage #photo-full{grid-column:1/-1;grid-row:1;height:65dvh}.viewer-prev,.viewer-next{grid-row:2;margin-top:8px}.viewer-prev{grid-column:1}.viewer-next{grid-column:3}#photo-caption{font-size:14px}.photo-viewer{max-height:96dvh;width:96vw}}
@media(max-width:480px){.nature-gallery{grid-template-columns:repeat(2,minmax(0,1fr))}}

/* Article selection, teaching and direct contact. */
.articles-section{background:var(--cream);scroll-margin-top:100px}
.articles-section h2,.training-section h2{font-size:clamp(36px,4vw,60px);margin-top:24px;line-height:1.12}
.article-grid{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:48px 5vw}
.article-card{border-top:2px solid var(--pine);padding-top:24px;min-width:0;display:flex;flex-direction:column;align-items:flex-start}
.article-meta{display:flex;flex-wrap:wrap;justify-content:space-between;gap:12px 24px;width:100%;font-size:13px;line-height:1.6;color:#52644e}
.article-meta>span{font-weight:700;font-size:11px;letter-spacing:.075em}
.article-card h3{font:600 clamp(24px,2.3vw,34px)/1.3 Manrope,sans-serif;letter-spacing:-.04em;margin:23px 0 18px}
.article-card h3 a{text-decoration:none;display:block}
.article-card h3 a:hover{text-decoration:underline;text-decoration-thickness:1px;text-underline-offset:5px}
.article-card h3 a:focus-visible{outline:3px solid #557565;outline-offset:6px}
.article-card h3 span{font-size:.75em;white-space:nowrap}
.article-card p{font-size:18px;line-height:1.7;color:#435747;margin:0 0 24px}
.article-source{margin-top:auto;font-size:13px;line-height:1.6;color:#52644e}
.article-archive{margin-top:40px;display:inline-flex;gap:20px;align-items:center}
.training-section{padding-top:75px;padding-bottom:80px;background:var(--cream);scroll-margin-top:100px}
.training-grid{display:grid;grid-template-columns:repeat(3,minmax(0,1fr));gap:3vw}
.training-grid article{border-top:1px solid #aabb9e;padding-top:24px;min-width:0}
.training-number{font:500 16px Manrope,sans-serif;color:#536f42}
.training-grid h3{font:600 clamp(24px,2.2vw,33px)/1.3 Manrope,sans-serif;letter-spacing:-.04em;margin:22px 0 16px}
.training-grid p{font-size:18px;line-height:1.7;color:#435747;margin:0}
.contact-actions{display:flex;align-items:center;flex-wrap:wrap;gap:20px;margin-top:28px}
.contact-actions .button{margin-top:0}
.contact-phone{font-size:18px;font-variant-numeric:tabular-nums;white-space:nowrap}
.contact-layout .contact-actions{margin-bottom:28px}
@media(max-width:900px){.training-grid{grid-template-columns:1fr;gap:32px}.training-grid h3{margin-top:15px}.training-grid p{max-width:650px}}
@media(max-width:720px){.article-grid{grid-template-columns:1fr;gap:36px}.articles-section .section-heading,.training-section .section-heading{display:block}.articles-section .section-heading p,.training-section .section-heading p{margin-top:25px}.article-card h3{font-size:27px}.article-card p,.training-grid p{font-size:17px}.article-archive{font-size:12px;gap:14px}.training-section{padding:55px 24px}.contact-actions{align-items:flex-start;flex-direction:column;gap:15px}.contact-actions .button{font-size:12px;gap:18px}.article-meta{font-size:12px}}